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<article language="en">
	<journal>
		<journal_title>Hydrology and Earth System Sciences</journal_title>
		<journal_url>www.hydrol-earth-syst-sci.net</journal_url>
		<issn>1027-5606</issn>
		<eissn>1607-7938</eissn>
		<volume_number>11</volume_number>
		<issue_number>6</issue_number>
		<publication_year>2007</publication_year>
	</journal>
	<doi>10.5194/hess-11-1717-2007</doi>
	<article_url>http://www.hydrol-earth-syst-sci.net/11/1717/2007/</article_url>
	<abstract_html>http://www.hydrol-earth-syst-sci.net/11/1717/2007/hess-11-1717-2007.html</abstract_html>
	<fulltext_pdf>http://www.hydrol-earth-syst-sci.net/11/1717/2007/hess-11-1717-2007.pdf</fulltext_pdf>
	<start_page>1717</start_page>
	<end_page>1730</end_page>
	<publication_date>2007-11-01</publication_date>
	<article_title content_type="html">Eco-geomorphology of banded vegetation patterns in arid and semi-arid regions</article_title>
	<authors>
		<author numeration="1" affiliations="1">
			<name>P. M. Saco</name>
			<email>patricia.saco@newcastle.edu.au</email>
		</author>
		<author numeration="2" affiliations="1">
			<name>G. R. Willgoose</name>
		</author>
		<author numeration="3" affiliations="2">
			<name>G. R. Hancock</name>
		</author>
	</authors>
	<affiliations>
		<affiliation numeration="1" content_type="html">School of Engineering, The University of Newcastle, Callaghan, New South Wales, 2308, Australia</affiliation>
		<affiliation numeration="2" content_type="html">School of Environmental and Life Sciences, The University of Newcastle, Callaghan, New South Wales, 2308, Australia</affiliation>
	</affiliations>
	<abstract content_type="html">The interaction between vegetation and hydrologic processes is particularly
tight in water-limited environments where a positive-feedback links soil
moisture and vegetation. The vegetation of these systems is commonly
patterned, that is, arranged in a two phase mosaic composed of patches with
high biomass cover interspersed within a low-cover or bare soil component.
These patterns are strongly linked to the redistribution of runoff and
resources from source areas (bare patches) to sink areas (vegetation
patches) and play an important role in controlling erosion.

&lt;br&gt;&lt;br&gt;

In this paper, the dynamics of these systems is investigated using a new
modeling framework that couples landform and vegetation evolution,
explicitly accounting for the dynamics of runon-runoff areas. The objective
of this study is to analyze water-limited systems on hillslopes with mild
slopes, in which overland flow occurs predominantly in only one direction
and vegetation displays a banded pattern. Our simulations reproduce bands
that can be either stationary or upstream migrating depending on the
magnitude of the runoff-induced seed dispersal. We also found that
stationary banded systems redistribute sediment so that a stepped
microtopography is developed. The modelling results are the first to
incorporate the effects of runoff redistribution and variable infiltration
rates on the development of both the vegetation patterns and
microtopography. The microtopography for stationary bands is characterized
by bare soil on the lower gradient areas and vegetation on steeper gradients
areas. For the case of migrating vegetation bands the model generates
hillslope profiles with planar topography. The success at generating not
only the observed patterns of vegetation, but also patterns of runoff and
sediment redistribution suggests that the hydrologic and erosion mechanisms
represented in the model are correctly capturing some of the key processes
driving these ecosystems.</abstract>
	<references>
		<reference numeration="1" content_type="text"> Ahnert, F.: The role of the equilibrium concept in the interpretation of landforms of fluvial erosion and deposition, in: L&apos;evolution des versants, edited by: Macar, P., p. 23&amp;ndash;41, University of Liege, Liege, 1967. </reference>
		<reference numeration="2" content_type="text"> Aguiar, M. R. and Sala, O. E.: Patch structure, dynamics and implications for the functioning of arid ecosystems, Trends in Ecology and Evolution, 14, 273&amp;ndash;277, 1999. </reference>
		<reference numeration="3" content_type="text"> Berg, S. S. and Dunkerley, D. L.: Patterned Mulga near Alice Springs, central Australia, and the potential threat of firewood collection on this vegetation community, J. Arid Environ., 59, 313&amp;ndash;350, 2004. </reference>
		<reference numeration="4" content_type="text"> Bergkamp, G., Cerda, A., and Imeson, A. C.: Magnitude-frequency analysis of water redistribution along a climate gradient in Spain, Catena, 37, 129&amp;ndash;146, 1999. </reference>
		<reference numeration="5" content_type="text"> Bhark, E. W. and Small, E. E.: Association between plant canopies and the spatial patterns of infiltration in shrubland and grassland of the Chihuahuan Desert, New Mexico, Ecosystems, 6, 185&amp;ndash;196, 2003. </reference>
		<reference numeration="6" content_type="text"> Bochet, E., Poesen, J., and Rubio, J. L.: Mound development as an interaction of individual plants with soil, water erosion and sedimentation processes on slopes, Earth Surf. Proc. Land., 25, 847&amp;ndash;867, 2000. </reference>
		<reference numeration="7" content_type="text"> Boer, M. and Puigdefábregas, J.: Effects of spatially structured vegetation patterns on hillslope erosion in a semiarid Mediterranean environment: a simulation study, Effects of vegetation patterns on erosion, Earth Surf. Processes Landforms, 30, 149&amp;ndash;167, 2005. </reference>
		<reference numeration="8" content_type="text"> Brown, J. H., Valone, T. J., and Curtin, C. G.: Reorganization of an arid ecosystem in response to recent climate change, Proceedings of the National Academy of Sciences (USA), 94, 9729&amp;ndash;9733, 1997. </reference>
		<reference numeration="9" content_type="text"> Cammeraat, L. H. and Imeson, A. C.: The evolution and significance of soil-vegetation patterns following land abandonment and fire in Spain, Catena, 37(1&amp;ndash;2), 107&amp;ndash;127, 1999. </reference>
		<reference numeration="10" content_type="text"> Cerdà, A.: Soil aggregate stability under different Mediterranean vegetation types, Catena, 32, 73&amp;ndash;86, 1998. </reference>
		<reference numeration="11" content_type="text"> Collins, D. B. G., Bras, R. L., and Tucker, G. E.: Modeling the effects of vegetation-erosion coupling on landscape evolution, J. Geophys. Res., 109, F03004, doi:10.1029/2003JF000028, 2004. </reference>
		<reference numeration="12" content_type="text"> d&apos;Herbes, J. M, Valentin, C., Tongway, D., and Leprun, J. C.: Banded Vegetation Patterns and related Structures, in: Banded vegetation patterning in arid and semiarid environments: ecological processes and consequences for management, Ecological studies 149, Springer-Verlag, New York, USA, 1&amp;ndash;19, 2001. </reference>
		<reference numeration="13" content_type="text"> Dunkerley, D. L.: Banded vegetation: development under uniform rainfall from a simple cellular automation model, Plant Ecology, 129, 103&amp;ndash;111, 1997. </reference>
		<reference numeration="14" content_type="text"> Dunkerley, D. L.: Assessing the influence of shrubs and their interspaces on enhancing infiltration in an arid Australian shrubland, Rangeland Journal, 22(1), 58&amp;ndash;71, 2000. </reference>
		<reference numeration="15" content_type="text"> Dunkerley, D. L.: Infiltration rates and soil moisture in a groved Mulga community near Alice Springs, arid central Australia: evidence for complex internal rainwater redistribution in a runoff-runon landscape, J. Arid Environ., 51, 199&amp;ndash;219, 2002. </reference>
		<reference numeration="16" content_type="text"> Dunkerley, D. L. and Brown, K. J.: Runoff and runon areas in a patterned chenopod shrubland, arid western New South Wales, Australia: characteristics and origin, J. Arid Environ., 30, 41&amp;ndash;55, 1995. </reference>
		<reference numeration="17" content_type="text"> Dunkerley, D. L. and Brown, K. J.: Banded vegetation near Broken Hill, Australia: significance of surface roughness and soil physical properties, Catena, 37, 75&amp;ndash;88, 1999. </reference>
		<reference numeration="18" content_type="text"> Dunkerley, D. L. and Brown, K. J.: Oblique vegetation banding in the Australian arid zone: implications for theories of pattern evolution and maintenance, J. Arid Environ., 51, 163&amp;ndash;181, 2002. </reference>
		<reference numeration="19" content_type="text"> Dunne, T., Zhang, W., and Aubry, B. F.: Effects of rainfall, vegetation, and microtopography on infiltration and runoff, Water Resou. Res., 9, 2271&amp;ndash;2285, 1991. </reference>
		<reference numeration="20" content_type="text"> Eagleson, P. S.: Dynamic Hydrology, McGraw-Hill Book Company, New York, 351 p., 1970. </reference>
		<reference numeration="21" content_type="text"> Eldridge, D. J. and Rosentreter, R. R.: Shrub mounds enhance water flow in a shrub-steppe community in southwestern Idaho, USA, in: Seed and soil dynamics in shrubland ecosystems, edited by: Hild, A. L., Shaw, N. L., Meyer, S., Booth, D. T., and McArthur, E. D., USDA Forest Service Proceedings RMRS-P-31, Ogden UT:US, 2004. </reference>
		<reference numeration="22" content_type="text"> Flanagan, D. C. and Nearing, M. A. (Eds.): WEPP: USDA-Water Erosion Prediction Project, NSERL Rep. 10, National Soil Erosion Lab., U.S. Dep. of Agric., Laffayette, Indiana, 1995. </reference>
		<reference numeration="23" content_type="text"> Fox, D. M., Bryan, R. B., and Price, A. G.: The influence of slope angle on infiltration rate and surface seal characteristics for interrill conditions, Geoderma, 80, 181&amp;ndash;194, 1997. </reference>
		<reference numeration="24" content_type="text"> Fox, D. M., Le Bissonnais, Y., and Bruand, A.: The effect of ponding depth on infiltration in a crusted surface depression, Catena, 32(2), 87&amp;ndash;100, 1998. </reference>
		<reference numeration="25" content_type="text"> Gao, Q. and Reynolds, J. F.: Historical shrub-grass transitions in the northern Chihuahuan Desert: modeling the effects of shifting rainfall seasonality and event size over a landscape gradient, Global Change Biology, 9, 1&amp;ndash;19, 2003. </reference>
		<reference numeration="26" content_type="text"> Gilad, E., von Hardenberg, J., Provenzale, A., Schachak, M., and Meron, E.: Ecosystem Engineers: From Pattern Formation to Habitat Creation, Phys. Rev. Lett., 93(9), 098105, doi:10.1103/PhysRevLett.93.098105, 2004. </reference>
		<reference numeration="27" content_type="text"> Henderson, F. M. and Wooding, R. A.: Overland flow and groundwater flow from a steady rainfall of finite duration, J. Geophys. Res., 69, 1531&amp;ndash;1540, 1964. </reference>
		<reference numeration="28" content_type="text"> HilleRisLambers, R., Rietkerk, M., van den Bosch, F., Prins, H. H. T., and de Kroon, H.: Vegetation pattern formation in semi-arid grazing systems, Ecology, 82, 50&amp;ndash;61, 2001. </reference>
		<reference numeration="29" content_type="text"> Imeson, A. C. and Prinsen, H. A. M.: Vegetation patterns as biological indicators for identifying runoff and sediment source and sink areas for semi-arid landscapes in Spain, Agriculture, Ecosyst. Environ., 104, 333&amp;ndash;342, 2004. </reference>
		<reference numeration="30" content_type="text"> Istanbulluoglu, E. and Bras, R. L.: Vegetation-modulated landscape evolution: Effects of vegetation on landscape processes, drainage density, and topography, J. Geophys. Res., 110, F02012, doi:10.1029/2004JF000249, 2005. </reference>
		<reference numeration="31" content_type="text"> Julien, P. Y., Saghafian, B., and Ogden, F. L.: Raster-based hydrologic modeling of spatially varied surface runoff, Water Resour. Bull., 31(3), 523&amp;ndash;536, 1995. </reference>
		<reference numeration="32" content_type="text"> Klausmeier, C. A.: Regular and irregular patterns in semiarid vegetation, Science, 284, 1826&amp;ndash;1828, 1999. </reference>
		<reference numeration="33" content_type="text"> Lavee, H., Imeson, A. C., and Sarah, P.: The impact of climate change on geomorphology and desertification along a Mediterranean-arid transect, Land Degradation and Development, 9, 407&amp;ndash;422, 1998. </reference>
		<reference numeration="34" content_type="text"> Lefever, R. and Lejeune, O.: On the origin of tiger bush, B. Math. Biol., 59, 263&amp;ndash;294, 1997. </reference>
		<reference numeration="35" content_type="text"> Ludwig, J. A., Wilcox, B. P., Breshears, D. D., Tongway, D. J., and Imeson, A. C.: Vegetation patches and runoff-erosion as interacting ecohydrological processes in semiarid landscapes, Ecology, 86(2), 288&amp;ndash;297, 2005. </reference>
		<reference numeration="36" content_type="text"> Ludwig, J. A., Tongway, D. J., and Marsden, S. G.: Stripes, strands or stipples: modelling the influence of three landscape banding patterns on resource capture and productivity in semi-arid woodlands, Australia, Catena, 37, 257&amp;ndash;273, 1999. </reference>
		<reference numeration="37" content_type="text"> Ludwig, J. A., Tongway, D. J., Bastin, G., and James, C.: Monitoring ecological indicators of rangeland functional integrity and their relation to biodiversity at local to regional scales, Austral Ecology, 29, 108&amp;ndash;120, 2004. </reference>
		<reference numeration="38" content_type="text"> Mitas, L. and Mitasova, H.: Distributed soil erosion simulation for effective erosion prevention, Water Resour. Res., 34(3), 505&amp;ndash;516, 1998. </reference>
		<reference numeration="39" content_type="text"> Mabbutt, J. A. and Fanning, P. C.: Vegetation banding in arid Western Australia, J. Arid Environ., 12, 41&amp;ndash;59, 1987. </reference>
		<reference numeration="40" content_type="text"> Noy-Meir, I.: Desert ecosystems: environment and producers, Ann. Rev. Ecol. Systematics, 4, 25&amp;ndash;51, 1973. </reference>
		<reference numeration="41" content_type="text"> Noy-Meir, I.: Spatial effects in modelling of arid ecosystems, in: Arid Land Ecosystems, edited by: Goodall, D. W. and Perry, R. A., pp 411&amp;ndash;433, Cambridge University Press, 1981. </reference>
		<reference numeration="42" content_type="text"> Porporato, A., Laio, F., Ridolfi, L., Caylor, K. K., and Rodriguez-Iturbe, I.: Soil moisture and plant stress dynamics along the Kalahari precipitation gradient, J. Geophys. Res., 108(D3), 4127, doi:10.1029/2002JD002448, 2003. </reference>
		<reference numeration="43" content_type="text"> Puigdefabregas, J. and Sanchez, G.: Geomorphological implications of vegetation patchiness in semi-arid slopes, in: Advances in Hillslope Processes, 2: 1027&amp;ndash;1060, edited by: Anderson, M. and Brooks, S., John Wiley, 1996. </reference>
		<reference numeration="44" content_type="text"> Puigdefábregas, J., Sole, A., Gutierrez, L., del Barrio. G., and Boer, M.: Scales and processes of water and sediment redistribution in drylands: results from the Rambla Honda field site in Southeast Spain, Earth-Sci. Rev., 48, 39&amp;ndash;70, 1999. </reference>
		<reference numeration="45" content_type="text"> Saco, P. M. and Willgoose G. R.: Modeling Complex Interactions between Ecology, Hydrology and Soil Erosion Patterns in Arid regions, EOS Trans. AGU, 87(52) Fall Meet. Suppl., Abstract H11G-02, 2006. </reference>
		<reference numeration="46" content_type="text"> Sparrow, A. D., Friedel, M. F., and Stafford Smith, D. M.: A landscape-scale model of shrub and herbage dynamics in Central Australia, validated by satellite data, Ecological Modelling, 97, 197&amp;ndash;216, 1997. </reference>
		<reference numeration="47" content_type="text"> Slatyer, R. O.: Methodology of a water balance study conducted on a desert woodland Acacia aneura community, Arid Zone Res., 16, 15&amp;ndash;26, 1961. </reference>
		<reference numeration="48" content_type="text"> Rietkerk, M., Boerlijst, M. C., Van Langevelde, F., HilleRisLambers, R., van de Koppel, J., Kumar, L., Prins, H. H. T., and de Rooam, A. M.: Self-organization of vegetation in arid ecosystems, American Naturalist, 160, 524&amp;ndash;530, 2002. </reference>
		<reference numeration="49" content_type="text"> Thiery, J. M., D&apos;Herbes, J. M., and Valentin, C.: A model simulating the genesis of banded vegetation patterns in Niger, J. Ecol., 83, 497&amp;ndash;507, 1995. </reference>
		<reference numeration="50" content_type="text"> Tongway, D. J. and Ludwig, J. A.: Vegetation and soil patterning in semi-arid mulga lands of Eastern Australia, Aust. J. Ecol., 15, 23&amp;ndash;34, 1990. </reference>
		<reference numeration="51" content_type="text"> Tongway, D. J. and Ludwig, J. A.: The conservation of water and nutrients within landscapes, in: Landscape Ecology, Function and Management: Principles from Australia&apos;s Rangelands, Chap. 2, edited by: Ludwig, J., Tongway, D., Freudenberger, D., Noble, J., and Hodgkinson, K., CSIRO Publishing, Melbourne, 13&amp;ndash;22, 1997. </reference>
		<reference numeration="52" content_type="text"> Tongway, D. J. and Ludwig, J. A.: Theories on the origins, maintenance, dynamics, and functioning of banded landscapes, in Banded vegetation patterning in arid and semiarid environments: ecological processes and consequences for management, Ecological studies 149, Springer-Verlag, New York, USA, 20&amp;ndash;31, 2001. </reference>
		<reference numeration="53" content_type="text"> Tongway, D. J., Ludwig, J. A., and Whitford, W. G.: Mulga log mounds: fertile patches in the semi-arid woodlands of eastern Australia, Aust. J. Ecol., 14, 263&amp;ndash;268, 1989. </reference>
		<reference numeration="54" content_type="text"> Valentin, C. and d&apos;Herbes, J. M.: Niger tiger bush as a natural water harvesting system, Catena, 37, 231&amp;ndash;256, 1999. </reference>
		<reference numeration="55" content_type="text"> Valentin, C., d&apos;Herbes, J. M., and Poesen, J.: Soil and water components of banded vegetation patterns, Catena, 37, 1&amp;ndash;24, 1999. </reference>
		<reference numeration="56" content_type="text"> Vieux, B. E.: Geographic information systems and non-point source water quality and quantity modelling, Hydrol. Processes, 5(1), 101&amp;ndash;113, 1991. </reference>
		<reference numeration="57" content_type="text"> Wakelin-King, G. A.: Banded mosaic (&quot;tiger-bush&quot;) and sheetflow plains: a regional mapping approach, Aust. J. Earth Sci., 46, 53&amp;ndash;58, 1999. </reference>
		<reference numeration="58" content_type="text"> Walker, B. H., Ludwig, D., Holling, C. S., and Peterman, R. M.: Stability of semi-arid savanna grazing systems, J. Ecol., 69, 473&amp;ndash;498, 1981. </reference>
		<reference numeration="59" content_type="text"> Wilcox, B. P., Breshears, D. D., and Allen, C. D.: Ecohydrology of a resource-conserving semiarid woodland: effects of scale and disturbance, Ecological Monographs, 73(2), 223&amp;ndash;239, 2003. </reference>
		<reference numeration="60" content_type="text"> Willgoose, G. R.: User Manual for EAMS SIBERIA 8.30, www.telluricresearch.com/siberia_8.30_manual.pdf, 2004. </reference>
		<reference numeration="61" content_type="text"> Willgoose, G. R., Bras, R. L., and Rodriguez-Iturbe I.: A physically based coupled network growth and hillslope evolution model: 1 Theory, Water Resour. Res., 27(7), 1671&amp;ndash;1684, 1991. </reference>
		<reference numeration="62" content_type="text"> Woolhiser, D. A. and Liggett, J. A.: One-dimensional flow over a plane: The rising hydrograph, Water Resour. Res., 3(3), 753&amp;ndash;771, 1967. </reference>
		<reference numeration="63" content_type="text"> Zhang, H.: Organic matter incorporation affects mechanical properties of soil aggregates, Soil Tillage Res., 31, 263&amp;ndash;275, 1994. </reference>
	</references>
</article>

