Abstract. Seasonal hydrologic forecasts derive their skill from knowledge of initial hydrologic conditions and climate forecast skill associated with seasonal climate outlooks. Depending on the type of hydrological regime and the season, the relative contributions of initial hydrologic conditions and climate forecast skill to seasonal hydrologic forecast skill vary. We seek to quantify these contributions on a relative basis across the Conterminous United States. We constructed two experiments – Ensemble Streamflow Prediction and reverse-Ensemble Streamflow Prediction – to partition the contributions of the initial hydrologic conditions and climate forecast skill to overall forecast skill. In ensemble streamflow prediction (first experiment) hydrologic forecast skill is derived solely from knowledge of initial hydrologic conditions, whereas in reverse-ensemble streamflow prediction (second experiment), it is derived solely from atmospheric forcings (i.e. perfect climate forecast skill). Using the ratios of root mean square error in predicting cumulative runoff and mean monthly soil moisture of each experiment, we identify the variability of the relative contributions of the initial hydrologic conditions and climate forecast skill spatially throughout the year. We conclude that the initial hydrologic conditions generally have the strongest influence on the prediction of cumulative runoff and soil moisture at lead-1 (first month of the forecast period), beyond which climate forecast skill starts to have greater influence. Improvement in climate forecast skill alone will lead to better seasonal hydrologic forecast skill in most parts of the Northeastern and Southeastern US throughout the year and in the Western US mainly during fall and winter months; whereas improvement in knowledge of the initial hydrologic conditions can potentially improve skill most in the Western US during spring and summer months. We also observed that at a short lead time (i.e. lead-1) contribution of the initial hydrologic conditions in soil moisture forecasts is more extensive than in cumulative runoff forecasts across the Conterminous US.