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HESS | Articles | Volume 24, issue 6
Hydrol. Earth Syst. Sci., 24, 3271–3288, 2020
https://doi.org/10.5194/hess-24-3271-2020
© Author(s) 2020. This work is distributed under
the Creative Commons Attribution 4.0 License.

Special issue: Linking landscape organisation and hydrological functioning:...

Hydrol. Earth Syst. Sci., 24, 3271–3288, 2020
https://doi.org/10.5194/hess-24-3271-2020
© Author(s) 2020. This work is distributed under
the Creative Commons Attribution 4.0 License.

Research article 25 Jun 2020

Research article | 25 Jun 2020

Field observations of soil hydrological flow path evolution over 10 millennia

Field observations of soil hydrological flow path evolution over 10 millennia
Anne Hartmann1, Ekaterina Semenova2, Markus Weiler2, and Theresa Blume1 Anne Hartmann et al.
  • 1Section Hydrology, GFZ German Research Centre for Geosciences, Potsdam, Germany
  • 2Chair of Hydrology, University of Freiburg, Freiburg, Germany

Correspondence: Anne Hartmann (aha@gfz-potsdam.de)

Abstract
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Preferential flow strongly controls water flow and transport in soils. It is ubiquitous but difficult to characterize and predict. This study addresses the occurrence and the evolution of preferential flow during the evolution of landscapes and here specifically during the evolution of hillslopes. We targeted a chronosequence of glacial moraines in the Swiss Alps to investigate how water flow paths evolve along with the soil-forming processes. Dye tracer irrigation experiments with a Brilliant Blue FCF solution (4 g L−1) were conducted on four moraines of different ages (30, 160, 3000, and 10 000 years). At each moraine, three dye tracer experiments were conducted on plots of 1.5 m ×1.0 m. The three plots at each moraine were characterized by different vegetation complexities (low, medium, and high). Each plot was further divided into three equal subplots for the application of three different irrigation amounts (20, 40, and 60 mm) with an average irrigation intensity of 20 mm h−1. The day after the experiment five vertical soil sections were excavated, and the stained flow paths were photographed. Digital image analysis was used to derive average infiltration depths and flow path characteristics such as the volume and surface density of the dye patterns. Based on the volume density, the observed dye patterns were assigned to specific flow type categories. The results show a significant change in the type of preferential flow paths along the chronosequence. The flow types change from a rather homogeneous matrix flow in coarse material with high conductivities and a sparse vegetation cover at the youngest moraine to a heterogeneous infiltration pattern at the medium-age moraines. Heterogeneous matrix and finger flow are dominant at these intermediate age classes. At the oldest moraine only macropore flow via root channels was observed in deeper parts of the soil, in combination with a very high water storage capacity of the organic top layer and low hydraulic conductivity of the deeper soil. In general, we found an increase in water storage with increasing age of the moraines, based on our observations of the reduction in infiltration depth as well as laboratory measurements of porosity. Preferential flow is, however, not only caused by macropores, but especially for the medium-age moraine, it seems to be mainly initiated by soil surface characteristics (vegetation patches and microtopography).

1 Introduction
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The ability of soil to store and to transport water is essential for its ecosystem services such as nutrient cycling or water and gas balances (Clothier et al.2008; Amundson et al.2015; Hatfield et al.2017; Shang et al.2018). Thus, the interaction of water and soil is an elementary foundation for the existence and functioning of terrestrial ecosystems. This interaction is part of a large network of interactions of various ecosystem components (flora, fauna, material and energy fluxes, geomorphological conditions, and climate), which are also necessary for the existence and functioning of ecosystems. Soil filters the percolating water, redistributes it to groundwater or stream water, or holds it against gravity and makes it available for plants.

The soil functions are influenced and controlled by soil properties, which can vary spatially on the small (Hu et al.2008) and large scales (vertically along the profile and horizontally across landscapes; Bevington et al.2016), as well as temporally. These properties include soil texture and structure, i.e., the pore and grain size distributions, which in turn control the storage and transport capacity of the soil. Additional factors influencing soil functions are climate, topography, and vegetation. In undisturbed natural systems these factors are usually assumed to be constant at the observational timescale, and the inherent system dynamics only become apparent on long timescales.

Preferential flow, which is defined according to Hendrickx and Flury (2001) as a phenomenon “where water and solutes move along certain pathways, while bypassing a fraction of the porous matrix”, has impacts on water storage (Rye and Smettem2017) and thus plant water availability. It furthermore affects the transport of nutrients and contaminants (Jarvis2007) throughout the vadose zone and consequently also soil chemistry (Jin and Brantley2011; Bundt et al.2000) and groundwater quality. Allaire et al. (2009) attribute rapid flow and mass transport to flow through earthworm burrows, cracks in soil, and flow paths resulting from soil layering and hydrophobicity. They defined four types of preferential flow: crack flow, burrow flow (created by soil fauna), finger flow, and lateral flow along layer interfaces, where flow in burrows and cracks is also often classified as macropore flow. We will in the following distinguish flow in macropores according to their origin as crack flow and biopore flow, where the latter includes channels by activities of roots and soil fauna. Preferential flow in the form of macropore flow occurs mostly in fine-textured soils, whereas finger and funnel flow rather occurs in soils with a coarse texture (Hendrickx and Flury2001). General factors which can cause preferential flow paths are surface structure and properties such as vegetation cover, microtopography or hydrophobicity, as well as subsurface soil properties such as soil structure and soil type, subsurface heterogeneities, flow instabilities, and plant root activities (Weiler and Naef2003; Clothier et al.2008; Bachmair et al.2009; Jarvis2007; van Schaik2009; Wang et al.2018). Soil water conditions were also found to have an influence on the preferential flow path characteristics (Gimbel et al.2016; Hardie et al.2011; Bogner et al.2008).

Many preferential flow-influencing properties such as soil structure, soil texture, or vegetation cover change during landscape evolution (e.g., Vilmundardóttir et al.2014; Egli et al.2010; Dümig et al.2011) and thus also lead to a change in the soil hydraulic behavior (Lohse and Dietrich2005), which in turn has a direct impact on the surface and subsurface water transport. We therefore assume that the age of the soil has an influence on the prevailing preferential flow paths and thus the type and also the depth extent of the preferential flow paths can change over time. Especially root activities can lead to the generation of preferential flow paths in deeper layers, which was found by Cheng et al. (2014) based on a comparison of young and older forest plantations. On a large timescale of several million years, Lohse and Dietrich (2005) found a transition from mainly vertical water transport in younger volcanic soils in the Hawaiian Islands to lateral water transport along the boundary of a subsurface clay layer. The younger soil was coarse textured with high saturated hydraulic conductivities along the profile and a rather low field capacity, whereas the older soil revealed a higher field capacity and a distinct reduction in saturated hydraulic conductivity throughout the profile due to clay accumulation. Yoshida and Troch (2016) observed a major change in flow paths from deep groundwater flow to shallow subsurface flow in volcanic catchments of ages between 200 000 and 82 million years. Thus, the change of major flow paths with time has already been studied at the timescale on the order of 100 000 to millions of years, but little is known about how flow paths change during these first 10 000 years of landscape development.

Areas with receding glaciers have been shown to be suitable for soil development studies (Crocker and Dickson1957; Douglass and Bockheim2006; He and Tang2008; Egli et al.2010; Dümig et al.2011; Vilmundardóttir et al.2014; D'Amico et al.2014). In the cool and humid climate regions of former glacial areas, the soils develop from mineral soils to soils with a highly organic topsoil. These organic soil types are less intensively studied with regard to their soil hydraulic behavior compared to mineral soils (Carey et al.2007). It is known that these soils differ in their soil hydraulic properties from mineral soils (high total porosity – up to 90 % – and a low bulk density; Carey et al.2007), but little is known about how this development impacts water flow paths. Therefore, this study addresses the occurrence and the evolution of preferential flow during the first 10 000 years of landscape evolution in glacial moraines in the Swiss Alps. More specifically, we test the hypotheses that (1) vertical subsurface flow path types and vertical extent of flow paths change through the millennia as (2) the proportion of macropore flow will increase due to the development of biopores, (3) the soil develops from a homogeneously mixed material into a depth-differentiated soil system, and (4) physical weathering leads to a reduction in particle size and an increase in porosity.

Dye tracer experiments and an analysis of soil texture and soil physical properties were used to investigate how water flow paths evolve with hillslope age. The hydropedological approach (Lin2003) that links pedon (Quisenberry et al.1993), landscape (Cammeraat and Kooijman2009), and hydrologic process studies has already been applied to the preferential flow phenomenon (Jarvis et al.2012). Dye tracer experiments combined with digital image processing have been applied successfully to study preferential infiltration in soils (Weiler2001; Bogner et al.2008; Blume et al.2008; Laine-Kaulio et al.2015; Hardie et al.2011; Cheng et al.2014). In our study we use this method to identify how flow paths change during the coevolution of soil, vegetation, and topography. Understanding the changes in preferential flow paths as a result of the natural coevolution of landscape-forming factors can provide valuable knowledge on how these systems can also change as a result of human intervention (Richter and Mobley2009).

2 Material and methods
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2.1 Study site

The study area is located in the foreland of the Stein glacier above the tree line in the central Swiss Alps, south of the Sustenpass in the Urner Alps (approximately 4743 N, 825 E). Elevations range from 1900 to 2100 m a.s.l. The area lies in the polymetamorphic Erstfeld gneiss zone, which is part of the Aar massif (Blass et al.2003). The geology is defined by metamorphosed pre-Mesozoic metagranitoids, amphibolites, and gneisses (Heikkinen and Fogelberg1980; Schimmelpfennig et al.2014); thus the material is mainly acidic and rich in silicate. The closest official weather station is located 18 km away at Grimsel Hospiz (4634 N, 819 E) at an elevation of 1980 m a.s.l. For the norm period from 1981 to 2010, the station recorded an annual mean temperature of 1.9 C and an annual precipitation of 1856 mm. The precipitation distribution throughout the year is fairly uniform with a slight increase in the winter months (Schweizerische Eidgenossenschaft2016). The glacier foreland consists of moraines with unconsolidated glacial till. The humid and cool climate together with the nutrient-poor substrate and a relative high water permeability of the glacial till favor the formation of podsolic soils and humus in this area (Heikkinen and Fogelberg1980).

The moraines of the Stein glacier were exposed due to its retreat to the south. Four moraines were selected for this study (see Fig. 1). Schimmelpfennig et al. (2014) conducted a detailed dating study of the Stein glacier moraines, based on high-sensitivity beryllium-10 moraine dating and found that the ages of three moraines range between 160 to 10 000 years. The age of a fourth moraine was dated to 30 years based on maps and aerial photos.

https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f01

Figure 1Location (left) and surface cover (right) of the four selected proglacial moraines of the Stein glacier. White circles show locations of one of the three Brilliant Blue experiment plots per age class. Photo of the location is provided by © Google (2020). Photos of the 30-, 160-, and 10 000-year-old moraines were taken after the Brilliant Blue experiment (photos taken by Florian Lustenberger).

The oldest moraine with an age of 10 000 years is facing northeast. The second oldest moraine with an age between 2000 and 3000 years is the only one facing south. The two youngest moraines were exposed in the years 1860 and 1980–1990 and thus have an age of 160 and 30 years, respectively. Both moraines are facing northeast and are located closer to the glacier tongue at a distance of approximately 1 km from the oldest moraines. Both moraines are south of the glacial lake Steinsee (1930 m a.s.l.), which is a proglacial lake that was formed by the glacier retreat in 1924 (Blass et al.2003).

The vegetation of the moraines was mapped in summer 2017 (Maier et al.2019). Pronounced differences in vegetation coverage and species distributions were found among the four age classes. The vegetation of the oldest moraine was dominated by a variety of prostrate shrubs together with small trees and several grasses. On the 3000-year-old moraine, a grassland cover with fern, mosses, sedges, and forbs was found. The two youngest moraines however showed a lower degree of vegetation complexity. On the 160-year-old moraine, a combination of grasses, lichen, forbs, and shrubs was present. The youngest moraine still shows only a sparse vegetation cover with mainly grass, moss, forbs, and a few shrubs.

2.2 Field experiments

The dye tracer experiments were conducted between mid July and mid August 2018. We used Brilliant Blue FCF as dye tracer due to its good visibility, high mobility, and non-toxicity. We used a concentration of 4 g L−1, at which the tracer shows good sorption and visibility (Weiler and Flühler2004). Three study plots were selected at each moraine, based on the degree of vegetation complexity (low, medium, and high complexity). Vegetation complexity is characterized by vegetation coverage, the number of species, and the plant functional diversity. The functional diversity is calculated based on specific leaf area, nitrogen content, leaf dry matter content, Raunkiær's life form, seed mass, clonal growth organ, root type, and growth form. The collection of the required data and calculation of the vegetation complexity was done by the geobotany group of the University of Freiburg and is described in more detail in Maier et al. (2019).

The size of each study plot was 1.5 m ×1.0 m. The distances between the three study plots at each moraine ranged from 10 to 100 m. Each plot was further divided into three equal subplots of 0.5 m ×1.0 m. Figure 2 shows the experimental design at each moraine and illustrates the irrigation procedure.

https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f02

Figure 2Illustration of the experimental design and soil sampling scheme at each moraine.

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The subplots were irrigated with three different amounts of dyed water (20, 40, and 60 mm) and an irrigation intensity of 20 mm h−1. The irrigation intensity is relatively high with a return period of 2.8 years (Fukutome et al.2017). In preparation of the tracer application, large vegetation in the form of shrubs and bushes was cut off to a height of a few centimeters to reduce interception. The tracer was applied with a hand-operated sprayer connected to a battery-powered pump which guaranteed a constant pressure for a uniform flow rate of 60 L h−1. For a time-efficient irrigation of the three subplots with three irrigation amounts, the irrigation procedure was divided into three steps. In the first step, all three subplots were irrigated simultaneously for 60 min in a sequence of 5 min of irrigation and a 5 min break. This provides an application of 20 mm to all three subplots. After finishing the first step the first subplot was covered to avoid any additional water input. In a second step, the other two subplots were simultaneously irrigated for an additional 60 min in a sequence of 5 min of irrigation and 10 min breaks. This provides an application of an additional 20 mm to each of the two remaining subplots. In the last step, only the third subplot was irrigated for 60 min in a sequence of 2 min of irrigation and 10 min breaks, while the other two plots remained covered providing an additional 20 mm to this subplot. After the end of tracer application, the entire plot was covered to avoid any disturbance by natural rainfall.

The next day each subplot was excavated in up to five profiles of 7 to 10 cm. After the profile cuts were made with pickaxes, spades, and hand shovels, the profile walls were cleaned. Hanging roots were cut off, and rocks were not removed but made visible. The profiles of each subplot were photographed with a Panasonic Lumix DMC-FZ18 camera and at a resolution of 2248 pixels ×3264 pixels. A big umbrella was used to provide a uniform light distribution in the photographs and to avoid direct sunlight. A wooden frame for geometric correction and a grayscale (Kodak) attached to the frame (Fig. 3) for later color adjustment were included in the photographs. Since dye tracer experiments only provide snapshots of flow patterns at 24 h after the irrigation, we cannot exclude the possibility that initial preferential flow paths were obliterated by a later downward movement of the infiltration front. However, as the probability of this special case is relatively low, we assume that these snapshots are a viable basis for the comparison of characteristic flow patterns along the moraine ages.

https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f03

Figure 3Exemplary image analysis procedure for the 40 mm irrigated subplot at the 3000-year-old moraine. (a) Photograph of the vertical soil profile with a wooden frame and an attached grayscale. (b) Software-generated tricolor image of the photograph. (c) Manually corrected tricolor image. Blue indicates stained soil; white indicates unstained soil; and orange indicates rocks.

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2.3 Image analysis

The image analysis procedure by Weiler (2001) was used to generate tricolor images of the photographs showing stained and unstained areas (Fig. 3). A detailed description of the method can be found in Weiler and Flühler (2004). Instead of using the original IDL (Interactive Data Language) software package, a similar Python version was used. Basically, geometric correction, background subtraction, and color adjustment were carried out to correct differences in image illumination and changes in the spectral composition of daylight. The delineation of rocks and plants was done manually. In the resulting tricolor image, the horizontal and vertical length of a pixel correspond to 1 mm. Due to poor lighting conditions or a heterogeneous background color distribution in the soil caused by material transitions, small stones, or organic matter, the image analysis software was not able to recognize all large dye stains as coherent objects. Thus, a manual correction of the images using the photographs was necessary (see Fig. 3).

For a quantitative comparison of the dye patterns, the maximum infiltration depth, the volume density, and surface area density, as well as the stained path width, were calculated. These parameters are frequently used for an objective comparison and description of the dye patterns (Weiler and Flühler2004; Bachmair et al.2009; Laine-Kaulio et al.2015; Cheng et al.2014; Gimbel et al.2016; Laine-Kaulio et al.2015; Mooney and Morris2008; Öhrström et al.2002). Volume and surface area density are originally steorological parameters which are used to relate three-dimensional structures to measured two-dimensional parameters (Weibel1979). The volume density corresponds to the dye coverage and can be derived from one-dimensional information by calculating the fraction of stained pixels for each depth. The volume density profile is defined by the fraction of stained pixels per depth and is calculated as the average of all excavated profiles per plot. The surface area density in one dimension is calculated by using the intercept density, which describes the number of intercepts between stained and unstained pixels divided by the horizontal width of the soil profile. The profile of the surface area density describes the amount of intercepts per depth and is then also averaged over all photographed profiles per plot. Volume density provides no information about whether the stained area is the sum of many small fragments or a few large ones; thus the volume density alone should not be used to characterize flow patterns, and the surface area density should be used as a supplementary parameter. A high surface area density indicates a large number of small features.

Following the method described by Weiler (2001), the resulting dye patterns were next classified into flow type categories based on the proportions of three selected stained path width classes (stained path width <20, 20–200 mm, and >200 mm) relative to the volume density. The stained path width is equal to the horizontal extent of a stained flow path (Weiler2001). This classification method distinguishes between five flow types: (1) macropore flow with low interaction, (2) mixed macropore flow (low and high interaction), (3) macropore flow with high interaction, (4) heterogeneous matrix flow and finger flow, and (5) homogeneous matrix flow. Dye patterns which cannot be classified as one of these flow types are categorized as undefined. The classification method based on proportions of the classes of stained path width is illustrated in Fig. 4.

https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f04

Figure 4Ternary diagram for flow type classification based on the proportion of the three stained path width classes (after Weiler2001).

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This method is based on the assumption that the dye patterns are mainly controlled by certain preferential flow processes and that each flow process creates a characteristic dye pattern that can be described by the extent and distribution of stained features.

This method was proven to be suitable for the investigation of Weiler (2001) and Weiler and Flühler (2004) and was also used in a variety of additional studies (Bachmair et al.2009; Gimbel et al.2016; Mooney and Morris2008). However, an extension of the flow type categorization was needed for our study site with soils of different ages, texture, and additionally a high stone content. In the extended classification, we avoid a clear differentiation between macropore flow and finger flow. The original classification assigns finger flow only when both classes of the medium-sized stained path width (20–200 mm) and the biggest stained path width (>200 mm) account for approximately half of the dye coverage. This implies that finger flow is only prevalent when the dye pattern is characterized by a majority of broader stained path widths, ignoring the fact that the size of the finger-like flow paths can vary over a broad range (Wang et al.2018). Thus, we argue that while finger flow and macropore flow are caused by different properties and flow mechanisms, they can lead to similar dye patterns and distributions of stained path width classes. This is especially the case for macropore flow with high interaction, which creates broader stained paths that could also be assigned to finger flow. Therefore, both flow types were considered in the extended classification for this class. Furthermore, it was observed that the presence of rocks within the image analysis interrupts homogeneous blue stained areas and thus leads to smaller stained path widths. Using the original classification scheme on a soil profile with high stone content suggests a heterogeneous flow pattern, which can be classified as heterogeneous matrix flow, finger flow, or macropore flow depending on the abundance of rocks. Therefore, an additional class has been introduced, which is used when homogeneous matrix flow between rocks takes place. The classification rule for the additional flow type class is based on the proportion of blue-dye coverage of the available permeable matrix space (profile width minus the sum of stone widths per row). If at least 95 % of the permeable space is stained by blue dye, the flow type is classified as matrix flow between rocks.

2.4 Soil sampling and laboratory analysis

Soil samples were taken during August and September of 2018 close to each dye tracer plot. For grain size analysis, two disturbed bulk soil samples per depth were taken at 10, 30, and 50 cm depth at each plot. The total of 72 samples was analyzed in the laboratory between November 2018 and January 2019 by using a combination of dry sieving (grain sizes >0.063 mm) and sedimentation analysis (grain sizes <0.063 mm) with the hydrometer method (Casagrande1934). Organic matter removal was only possible by floating off the lighter fractions prior to grain size analysis. Since three plots were selected per moraine for the Brilliant Blue experiments, six samples per depth and age class (a total of 18 samples for each moraine) were available for the grain size analysis. Grain sizes between 2 and 0.063 mm were classified as sand, between 0.063 and 0.002 mm as silt, and smaller than 0.002 mm as clay. Grain size fractions of particles <2 mm were calculated as weight percentages of total weight of particles <2 mm, thus excluding gravel and stones to avoid single larger stones shifting or dominating the distribution. The gravel and stone fraction was calculated separately as a weight percentage of the entire soil sample.

For the analysis of the structural parameters, soil samples were taken with sample rings to provide undisturbed cores which preserve the natural soil structure. At each plot, two 250 cm3 and one 100 cm3 undisturbed soil samples were taken at a depth of 10 and 30 cm. Three samples of 100 cm3 were taken at 50 cm depth. Thus, per age class, nine samples per depth were available for the determination of the structural parameters of porosity and bulk density. A detailed overview of the sampling scheme at each plot is given in Fig. 2.

The porosity was determined in the lab using the water saturation method. For this method, sample weights were recorded at saturation and after drying at 105 C. For saturation, the samples were placed in a small basin. The water level in the basin was increased stepwise by 1 cm d−1. When the water level reached the top of the soil sample and the sample was fully saturated, the bottom of the sample was sealed, and the weight at saturation was measured. Bulk density was determined by relating the dry mass after drying at 105 C to the sample volume. The loss on ignition is a measure of the organic substance in the soil and describes the proportion of the organic substance that was oxidized during annealing for 24 h at 550 C. The loss on ignition was determined by drying subsamples (4–6 g) for at least 24 h at 105 C and then at 550 C. The ignition loss is then calculated by relating the weight loss after drying at 550 C to the sample weight after drying at 105 C.

2.5 Statistical analysis

The nonparametric Kruskal–Wallis test was used to test the significance of the differences in the soil texture among the four moraines of differing age classes. It can be applied when the assumption of a normal distribution cannot be made and is also valid for small sample sizes. We applied the test to each grain size fraction across the four age classes. Average values were based on 18 samples per age class. The grain size distribution at 10 cm depth of the oldest moraine was excluded from consideration, since due to the high organic content not all organic matter could be removed, and the results may therefore be erroneous.

3 Results
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3.1 Soil texture and structural parameters

Comparing the depth-averaged soil texture over the millennia, we find that while the soil texture at the youngest moraine mainly consists of sand, the grain sizes decrease over the millennia with silt being the largest fraction after 10 000 years (Fig. 5). Clay content increased with age for the three older moraines, with the youngest moraine being the exception (having the second-highest clay content). The Kruskal–Wallis test with a 0.05 confidence level showed that differences in grain size fractions among the four age classes were statistically significant (p values <0.05; sand of p=0.0013, silt of p=0.0006, and clay of p=0.0018).

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Figure 5(a) Profile-averaged grain size fractions for the four moraines. Fractions are percentages of the fine earth fraction (<2 mm). (b) Profile-averaged gravel content (>2 mm) calculated as the percentage of the entire sample weight. Each average is based on 18 samples.

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A significant reduction in grain size over time was observed, which is most pronounced between 3000 and 10 000 years of soil development. The gravel and stone fraction is roughly the same at the three younger moraines and significantly lower at the oldest moraine (Fig. 5b). The structural parameters of porosity and bulk density also show a clear trend with age, with porosity increasing and bulk density decreasing (Fig. 6).

Table 1Linear rates of change in porosity, bulk density, and grain size between adjacent age classes calculated based on median values. Mass fraction: mf %.

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https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f06

Figure 6Evolution of soil porosity (a), bulk density (b), and loss on ignition (c) in 10, 30, and 50 cm depth.

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The porosity observed at the youngest moraine ranges between 0.22 and 0.37, with no pronounced differences among the three soil depths. The 160-year-old moraine has a higher porosity in the upper 10 cm than the 30-year-old moraine. After 3000 years the increase in porosity continues but is now also visible in 30 and 50 cm, but with much higher values at 10 cm (Fig. 6a). After 10 000 years the porosity at 10 cm ranges from 0.6 to up to more than 0.8. The other two depths also experienced a further increase in porosity. The decrease in bulk density is also most pronounced in the top layer of the soil (Fig. 6b). While after 30 years the bulk density in the upper 10 cm ranges around 1.7 g cm−3, the bulk density after 10 000 years is much smaller and ranges between 0.2 and 0.7 g cm−3. After 3000 years the trend is also visible at 30 and 50 cm.

The loss on ignition, as a measure for the organic matter content, shows an increase throughout the first 10 millennia of soil development, which is most pronounced in the upper soil layer (see Fig. 6c). At the two youngest moraines, the organic matter content is still very low (<2 wt %; percentage by weight). At these two age classes, the organic matter content is homogeneously distributed over the profile, with a slight tendency to higher values in the topsoil at the 160-year-old moraine. The 3000-year-old moraine shows a strong increase in the organic matter content in the surface layer. At the oldest moraine, the trend of increasing organic matter continues in all three depths. Here, the organic matter content in the topsoil makes up to two thirds of the soil material. However, the organic matter content varies distinctly with a minimum of 6 wt % and a maximum of 87 wt %. In deeper depths, the organic content also increases compared to the 3000-year-old soil but remains below 20 wt %.

Even though the differences in soil physical characteristics between 30 and 160 years are comparatively small, the rates of change during this initial phase are highest (Table 1). Between 160 and 3000 years, the rates of change are significantly reduced and remain in a similar range between 3000 and 10 000 years.

3.2 Vertical-dye pattern analysis

Flow patterns traced with Brilliant Blue dye changed considerably with moraine age (Fig. 7). The volume density profile is a measure of the amount of blue dye per depth. The profile patterns of volume and surface area density show distinct differences among age groups, while differences between the vegetation complexity levels are not as clear (Figs. 7 and 8).

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Figure 7Volume density profiles per age class, vegetation complexity, and irrigation amount. The volume density is the fraction of stained pixels, here colored by flow path width (stained path width; SPW) and rock sizes.

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https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f08

Figure 8Surface area density profiles per age class, vegetation complexity, and irrigation amount (20, 40, and 60 mm). A high surface area density indicates a large number of small features.

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The volume density of the blue dye was classified in three selected groups of stained path width (Weiler2001). Additionally, the volume density of rocks was also classified in three groups (Fig. 7).

An analysis of the average or maximum infiltration depth based on the dye profiles was not possible because not all profiles could be excavated up to the maximum infiltration depth. In most cases, large boulders prevented further excavation or the infiltration depth was more than 1 m. The latter was mostly the case at the youngest moraines. Only at the oldest moraine could a maximum infiltration depth be determined based on the dye profiles.

Combining the volume density profiles with the surface area density profiles, it is possible to derive whether the stained area described by the volume density is made up of many small flow paths or few large ones. The shapes of the volume density profiles and the surface area density of the youngest moraine are all very similar across the vegetation complexity levels and irrigation amounts. The youngest moraine has a higher volume density of flow paths in the top half of the soil profile than all other moraines (Fig. 7). There are almost no unstained areas. Beginning from approximately 30 cm depth, the volume density declines. The surface area density profiles show an opposite pattern (Fig. 8). The surface area density is smaller in the upper half and increases in the lower half of the profile. The combination of both parameters indicate a homogeneous staining in the top half, where interruptions of stained areas are only caused by rocks. In the lower part of the soil profiles the flow paths are subdivided, which is indicated by the increase in surface area density (apparent at the plots at 30 cm depth for low, 25 cm depth for medium, and 50 cm depth for high vegetation complexity). This combined with a decline in volume density indicates a narrowing of the flow paths. For the plots of low and high vegetation complexity, the change in flow paths coincides with a layer of higher clay and silt content. This layer does not exist at the plot of medium vegetation complexity. In this case, the narrowing and splitting up of flow paths is caused by large rocks. No clear differences are visible between the different irrigation amounts. The proportion of classes of the stained path width is controlled by the existence of rocks. The maximum infiltration depth is either controlled by the position of the clay layer or is deeper than the profile depth and therefore cannot be determined.

Comparing the 160-year-old moraine to the youngest moraine, we find that the volume density is lower and the surface density is higher in the upper part of the profile. Also unstained areas (colored white in Fig. 7) are visible, which indicates that the higher surface density is not caused by the existence of rocks that split up a homogeneously stained area as was the case for the youngest moraine. In this case there is no total dye coverage of the permeable soil, and the preferential flow paths are initialized already near or at the soil surface. The surface density profiles show a decrease in the lower half of the soil profile, which either goes along with a decrease in the blue-dye coverage and an increase in the stone coverage or an increase in the blue-dye coverage without a significant change in stone coverage. Both indicate a reduction in the amount of separate flow paths and an increase in flow path widths, even if the permeable space is reduced due to an increase in rock content. Also the fraction of stained path width (SPW) bigger than 200 mm increases, which indicates that the dye plumes widen in deeper soil depths.

Compared to the two youngest moraines, the moraine of 3000 years shows in general a higher surface area density and a lower dye coverage combined with a higher fraction of an unstained permeable soil matrix. This indicates that similar to the 160-year-old moraine, water is transported in individual flow paths, but here there are more flow paths, and they have a smaller width. This can also be seen in the less frequent appearance of stained path width higher than 200 mm (Fig. 7). Similar to the 160-year-old moraine, preferential flow paths are already initialized at the top of the soil during infiltration (apparent from the white-colored areas across the profile in Fig. 7).

The oldest moraine with an age of 10 000 years shows the highest surface area density and the lowest volume density combined with the lowest infiltration depths. The surface and volume density profiles show the same pattern: after a peak close to the soil surface, both density profiles show a decrease with soil depth. This means that the dyed water is only transported deeper into the soil via a few individual flow paths.

https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f09

Figure 9Profiles of flow types per age class, vegetation complexity, and irrigation amount.

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https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f10

Figure 10Relative frequency distribution of flow types of the four moraine age classes. Basically all observations fit into the six flow type categories. The fraction of observations categorized as “undefined flow type” is negligible.

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3.3 Flow type classification

Using the information in the volume density profiles and the stained path widths to characterize flow types (Weiler2001), we found a trend from a rather homogeneous flow pattern with matrix flow in a fast-draining coarse-textured soil at the youngest moraine to a more heterogeneous flow pattern with a mix of heterogeneous matrix flow and finger flow at both medium-age moraines (Fig. 9). While the flow characteristics of both the 160- and 3000-year-old moraine are dominated by finger flow with smaller stained path widths, this is much more pronounced in the 3000-year-old moraine. By contrast, the oldest moraine stands out clearly from the other age groups. Here, only the macropore flow is predominant. For the deeper soil layers, this was confirmed during the field experiments, since the few macropore flow paths were clearly visible. For the topsoil, the result is less certain, as the blue areas were very difficult to identify during the image analysis due to the very dark color of the organic layer.

The relative frequency distribution of the flow types per moraine age class derived from the results in Fig. 9 shows a clear shift in the flow type distribution along the age classes (Fig. 10). At the youngest moraine, all types of finger flow and matrix flow are present, and the frequency distribution does not show a distinct peak at any flow type. With increasing age, macropore flow becomes more and more important, and the peaks in the frequency distribution become more and more pronounced (Fig. 10).

4 Discussion
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4.1 Evolution of soil texture and structure

The observation of bulk density, porosity, and soil texture show significant differences between the age groups as well as some clear trends with age. A detailed description of this data set can be found in (Hartmann et al.2020).

4.1.1 The early years: 30–160

The 30-year-old soil is characterized by coarse material, with a soil texture composed of almost three quarters of sand. The soil texture observed at the 160-year-old moraine does not differ strongly from the 30-year-old moraine, whereas slight changes are already evident in porosity and bulk density. These findings are similar to findings by Dümig et al. (2011), who found no specific trend in grain size distribution for soils in the range of 15 to 140 years at a soil chronosequence in the foreland of the retreating Damma glacier (Switzerland). They furthermore found a high variability in grain size distribution within the single age classes. However, in the same study a slight decrease in variability with increasing age and a noticeable higher clay content were found at the reference site with an age of more than 700 years. He and Tang (2008) also revealed a nonlinear increase in the maximum clay content for soils up to 180 years at a glacier foreland in a monsoon temperate region in southwestern China.

After 160 years of soil development, the porosity in the top layer increased, and bulk density decreased. In general, these changes could be linked to changes in grain sizes, as the breakdown of particles leads to an increase in total pore space (porosity) and thus to a reduction in bulk density (Arvidsson1998). However, since changes in grain sizes were only marginal, the vegetation development, which includes an increase in the root activities, litter accumulation, and biological activities in the root zone, is likely the main cause for changes in bulk density and porosity (Neris et al.2012; Carey et al.2007). The vegetation coverage of both moraines differs significantly (Maier et al.2019). The youngest moraine still shows only low vegetation cover with only single plants (mainly grasses and forbs) and little root mass with an observed maximum rooting depth of 15 cm (occasionally up to 30 cm), whereas the 160-year-old moraine already has a relatively closed vegetation cover with a combination of shrubs and smaller plants like forbs and grasses forming a loose root network with roots up to a maximum diameter of 5–6 mm and a maximum depth of 35 cm (as observed during the excavation of the soil profiles).

Similar findings in bulk density evolution were also observed by Crocker and Major (1955), who found a decrease in the bulk density over the first 200 years of soil development from more than 1.4 g cm−3 to less than 0.8 g cm−3 for glacial till in southeastern Alaska. A less pronounced reduction was also found by Crocker and Dickson (1957). He and Tang (2008) found a reduction for the time span of 180 years from approximately 1.42 to 0.95 g cm−3 that was also more distinct in the upper horizon. Vilmundardóttir et al. (2014) revealed at a glacier foreland in southeastern Iceland under maritime climate conditions a reduction from 1.36 to 1.07 g cm−3 for a time span of 120 years. All studies mentioned above linked this decrease in bulk density to the vegetation development with time.

4.1.2 Intermediate stage: 3000 years

At 3000 years of soil development we observe a distinct increase in silt and a reduction in sand content, and this development continues, as observed in the 10 000-year moraine, where the silt content now makes up the largest share. These findings agree with findings by Douglass and Bockheim (2006), who studied several moraines in Buenos Aires with ages ranging from 16 000 years to 1 000 000 years and found an accumulation of clay-sized particles with increasing age, but with a decrease in the accumulation rate over the years. A high fraction of silt is very common for soils in mountain areas (Ellis1992). Physical weathering due to high fluctuations between day and night temperature and freezing cycles (Birse1980) leads to a reduction in grain size, without changing the particle mineralogy (Ellis1992).

The soil material at the 30-year-old moraine showed a relatively uniform porosity and bulk density throughout the profile. After 3000 years, porosity increases, and bulk density decreased even further, and this development is now also visible in deeper soil depths. The continuous increase in porosity and reduction in bulk density can be attributed to the continuing change in soil texture on the one hand and on the other hand to the pronounced vegetation development. Especially the latter, with the resulting accumulation of soil organic matter (see Fig. 6c) and the growth of an even denser root network that is now over 35 cm deep, is the main cause for the pronounced changes in the topsoil.

4.1.3 The late stage: 10 000 years

The oldest moraine shows a significantly higher silt content and porosity compared to the 3000-year-old moraine and a significantly lower bulk density. The change is visible at all soil depths, with the porosity in the uppermost depth being distinctly higher than the other depths. These differences in soil properties between the soil layers also indicate a progressive formation of distinct horizons in the soil.

The significantly higher porosity in the upper layer of the oldest moraine is caused by its thick organic layer (thickness up to 20 cm), which is characterized by porosity of up to 90 % (this was also found by Nyberg1995, in sandy–silty till on the western coast of Sweden and Carey et al.2007, in organic soils in a permafrost region in northwestern Canada).

Musso et al. (2019) investigated the evolution of pore sizes in the top 5 cm at the same soil chronosequence and found an increase in the number of small soil pores and a decrease in the relative proportion of macropores (pore diameter >0.05 mm) between 160 and 10 000 years. Thus the high porosity in the organic top layer at the oldest moraine is mainly composed of small pores. The top layer therefore has an increased water storage and water holding capacity. Due to the finer soil texture and higher porosity, the total water storage capacity of the oldest moraine is larger than that of the younger moraines.

An investigation of the saturated hydraulic conductivity evolution of the near surface (in 0–5, 5–20, and 20–40 cm) at the same chronosequence by Maier et al. (2019) found a decrease with increasing moraine age and soil depth. Saturated conductivity was found to be negatively correlated with the fraction of fine particles. The decrease in gravel content and the increase in silt seem to have an even a stronger effect on the saturated conductivity than the root network development (Maier et al.2019).

4.1.4 Soil heterogeneity and vegetation complexity

It is well known that soil properties are spatially heterogeneous (Bevington et al.2016; Hu et al.2008). As it was not possible to account for this variability with a large sample size, i.e., with a large number of experiments, we decided to take a different approach. Assuming that vegetation cover and subsurface flow paths are strongly linked, we took the variability in vegetation cover as a proxy and used it in an attempt to bracket this variability: per moraine three locations that differ in their vegetation complexity (low, medium, and high) were chosen for soil sampling and the dye tracer experiments. The analysis of the structural soil properties shows that there is a slight increase in spatial heterogeneity with age, especially in the topsoil (increase in the interquartile ranges for all properties in the top layer in Fig. 6), but occasionally also individual depths show a higher heterogeneity, irrespective of age.

The flow path analysis differentiated according to the vegetation complexity showed no systematic influence of the complexity level on the results. Heterogeneities within the individual experimental subplots were taken into account by averaging the volume density and surface area density across the five vertical profiles per subplot instead of relying on individual profiles. We therefore assume that the results of the flow path analysis are sufficiently representative to investigate their evolution across the chronosequence.

4.2 Evolution of flow paths

The flow type classification by Weiler (2001) was used to classify the volume density patterns into flow type categories. A comparison between the observations made during the excavation and the derived flow types showed that in this case an adaptation of the flow type classification was necessary. On the one hand, this adaptation involved the treatment of rocks to prevent misclassification, and, on the other hand, we introduced the possibility of finger-like flow paths with smaller widths. After these adaptations the derived flow types correspond well with the observations made in the field.

The observed staining patterns and derived flow types show a significant difference between the age groups, whereas no significant difference was observed with respect to vegetation complexity and irrigation amount.

4.2.1 The early years: 30–160

At the 30-year-old moraine, the water infiltrates homogeneously into the soil, probably due to the very low vegetation coverage and the coarse material texture. The dye pattern showed a mainly homogeneous staining of the soil material; thus derived flow types are mainly matrix flow in the form of homogeneous and heterogeneous matrix flow as well as matrix flow between rocks. Also finger flow occurs at the boundary to the clay layer or is caused by large blocks of rock, which are surrounded by clay. The determined macropore flow takes place only within the clay layer at a depth below 50 cm. In the clay layer, no significant biopores were identifiable, which is why it is assumed that the water is transported in cracks or along material interfaces. The upper coarse soil material with large pores and a low water holding capacity causes the water to be transported quickly deeper into the soil.

After 160 years, the derived predominant flow types shift to heterogeneous matrix flow and finger flow. The observed widening of the dye plumes in deeper soil depths might be caused by a change in material or a reduction of hydrophobicity with soil depth where the influence of plants and organic material decreases (Blume et al.2009). The dye coverage images show unstained soil areas starting also at the top of the soil profile, which indicates that preferential flow paths are initialized already at the soil surface or in the near-surface layer. This was also observed during the irrigation, where we saw that the irrigated water often mainly infiltrated in depressions. Grass patches also tend to inhibit infiltration. It was observed that in the presence of dense grass patches, the water infiltrated only next to the patches, leaving the area below the patches unstained.

4.2.2 Intermediate stage: 3000 years

Similar observations were made at the 3000-year-old moraine. The image analysis revealed that preferential flow paths start at the soil surface and thus are initiated by vegetation and microtopography causing a heterogeneous infiltration pattern. Field observations also revealed that heterogeneous infiltration was not only created due to dense grass patches but also occurred under relatively homogeneous grass cover. A laboratory test of the water drop penetration time (DeBano1981; Doerr et al.2000) on a soil sample of the upper soil material showed that the organic layer is highly water repellent in dry conditions (air dried for 2 weeks; water drop penetration time >10 min). An increase in the hydrophobicity index (Tillman et al.1989) with increasing moraine age was also found by Maier et al. (2019). Thus, we conclude that the hydrophobicity of the organic top layer has a big impact on infiltration and the initiation of unstable flow. Unstable flow occurs when horizontal wetting fronts break into fingers or preferential flow paths during the downward movement (Hendrickx and Flury2001). Compared to the 160-year-old moraine, the 3000-year-old moraine is characterized by a higher number of narrower preferential flow paths.

The derived dominant flow type class at the 3000-year-old moraine is macropore flow with high interaction and finger flow. Of both possible flow types, finger flow is the prevalent flow process causing the dye pattern. Several studies linked the formation of finger-like flow paths to hydrophobic properties of the soil (Wallach and Jortzick2008; Dekker and Ritsema2000; Ritsema and Dekker1994; Blume et al.2008; Wang et al.2018; Hardie et al.2011). It is assumed that hydrophobic compounds that are released during the decay of litter (Reeder and Jurgensen1979) or by root activity (Doerr et al.1998) coat soil particles or are deposited in the pore space and thus create a hydrophobic soil matrix (Doerr et al.2000). The humid and cool climate of former glacial areas leads to a slow decomposition of vegetation and thus to an accumulation of hydrophobic compounds (Doerr et al.2000).

4.2.3 The late stage: 10 000 years

At the oldest moraine, we saw a distinctly shallower infiltration depth (Fig. 7). During the experiment, no surface runoff was observed. Most of the water was stored in the organic top layer. The soil beneath the top layer was almost completely unstained, and water was transported only via a few macropores into deeper layers. A dense network of roots was only observed in the organic top layer, which included the thicker roots of the alpenrose (Rhododendron ferrugineum). The root network in the soil underneath was less dense with roots of smaller diameters but extended to a depth of more than 50 cm. Although the vegetation cover has been reduced to decrease interception, the interception storage capacity at the oldest moraine is still comparatively high. Thus, a reduction in the water available for infiltration cannot be ruled out.

4.2.4 Impact of rocks

The rock content at the 30-, 160-, and 3000-year-old moraines is relatively high, with especially large rocks (widths >20 mm) in deeper parts of the soil (Figs. 5b and 7). The large rocks lead to a reduction of the permeable area and thus can cause funnel flow (Hendrickx and Flury2001). This type of preferential flow was especially observed at the youngest moraine, where large boulders (>25 cm) located at deeper soil depths were surrounded by unstained fine-textured material. Smaller-sized rocks in the upper part seemed not to have an influence on water transport (apart from reducing the flow-through volume), since these rocks and the surrounding soil were completely stained.

A splitting of flow paths caused by rocks was also observed a few times at the 160- and 3000-year-old moraines. In this case, water flowing past the sides of medium-to-large-sized rocks creates a type of finger flow that is not caused by water repellency or air entrapment. The tendency of higher rock contents to increase the number of flow paths was also found by Bogner et al. (2014).

https://www.hydrol-earth-syst-sci.net/24/3271/2020/hess-24-3271-2020-f11

Figure 11Sequence of observed characteristic dye patterns and derived flow path controls compared to the qualitative evolution of soil texture, structural parameters, and vegetation. The shade of the root mass distribution triangles is a measure for the vertical root mass distribution with a darker color indicating a higher root mass. The width of the triangles is a measure for the root mass comparison between the moraine ages, with broader triangles indicating a higher root mass.

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4.2.5 Flow path controls along the age gradient

Integrating all of our findings on soil structural parameters, texture, vegetation cover, and flow path patterns provides an overview over their coevolution and highlights the derived major flow path controls (Fig. 11).

Along the coevolution of soil and vegetation over 10 000 years the major controls of subsurface flow paths change. At the youngest moraine flow paths are only controlled by soil texture. The coarse material leads to the deep downward movement of the infiltration front. Preferential flow paths only occur at the interfaces between coarse and finer material.

At the medium-age moraines, flow paths are mainly controlled by vegetation shielding, microtopography, and hydrophobicity. The latter is assumed to have an increased impact at the 3000-year-old moraine. After 10 000 years of hillslope evolution, subsurface water transport is highly preferential and controlled by flow paths caused by root channels or boundaries of textural classes. Water storage in the organic layer which is also the main rooting horizon increases strongly.

5 Conclusions
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Using Brilliant Blue dye experiments and soil sampling, we investigated the evolution of water transport paths along soil-forming processes. To our knowledge, this is the first study examining flow path evolution across the millennia in such detail. The evolution of the grain size distribution shows that grain size decreases with increasing age. The biggest changes are in the sand and silt fraction. Furthermore, water flow-defining structural parameters such as porosity and bulk density change during soil development, resulting in an increasing water storage capacity with age. The depth-dependent evolution of these parameters supports our hypothesis that the soil material develops with increasing age from a homogeneously mixed material to a depth-differentiated soil system with vertical gradients in flow and storage defining soil properties. Changes in these flow-defining parameters are caused by the evolution of grain size distribution and vegetation.

The derived flow types also support our hypothesis that vertical subsurface flow path types and their vertical extent change through the millennia. Flow types change from a homogeneous matrix flow in a fast-draining coarse-textured soil to a heterogeneous matrix and finger flow over the first 100–3000 years. At very young moraines, the water is homogeneously distributed within the soil matrix. However, the water storage capacity is relatively low due to the coarse material, and water is transported quickly deeper into the soil due to the high drainability. At the medium-age moraines, water is transported preferentially via finger-like flow paths deeper into the soil by leaving parts of the soil dry. With increasing hillslope age, we expected macropores induced by root activities to become more important. After 10 000 years, where the amount of soil matrix macropores decreased significantly, the macropore flow along roots plays an important role but is not very pronounced. Only a few roots reach beyond the organic top layer. However, this allows for a fast transport of water from the upper layer into deeper soil. The organic top layer has a pronounced influence on the soil water budget, by storing a significant amount of water. The increase in water storage with increasing age of the moraines also caused a reduction in infiltration depth.

The proportion of preferential flow paths increases with soil age. Preferential flow is, however, not only caused by macropores, but especially for the medium-age moraines it seems more controlled by soil surface characteristics such as vegetation patches, microtopography, and hydrophobicity. Thus, the evolution of flow paths is tightly linked to the complex interplay of soil-forming processes and vegetation development over the millennia. A lot of changes in vegetation cover, soil (hydraulic) properties, and flow paths occur within the first 160 years.

It was shown that the complex interaction of vegetation and soil development and its proven effect on flow path development also impacts the water balance, as the storage and conductivity properties of the soil change. However, the interplay between preferential flow paths and the soil hydraulic behavior not only influences the soil water budget but also runoff formation. These findings provide important insights into hydrological flow path evolution in transient systems.

Data availability
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Data availability. 

The soil physical data are described in detail and can be downloaded from (Hartmann et al.2020). The Brilliant Blue images and data are available upon request.

Author contributions
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Author contributions. 

AH and ES conducted the field experiments, prepared the images, and performed the analysis. AH collected the soil samples and conducted the laboratory analysis. MW and TB were involved in planning the fieldwork. AH prepared the paper with contributions from all coauthors.

Competing interests
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Competing interests. 

The authors declare that they have no conflict of interest.

Special issue statement
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Special issue statement. 

This article is part of the special issue “Linking landscape organisation and hydrological functioning: from hypotheses and observations to concepts, models and understanding (HESS/ESSD inter-journal SI)”. It is not associated with a conference.

Acknowledgements
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Acknowledgements. 

We thank Moritz Lesche and Jonas Freymüller for their assistance in the field. We also thank Kraftwerke Oberhasli AG (KWO) for permission to conduct the experiments. Many thanks to Thomas Michel and his team from the Alpin Center Sustenpass and Peter Luchs for their kind hospitality and support. We thank Nicholas Jarvis and one anonymous reviewer for their effort and improving our paper.

Financial support
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Financial support. 

This research has been supported by the German Research Foundation (DFG) and the Swiss National Science Foundation (SNF) within the DFG–SNF project HILLSCAPE (Hillslope Chronosequence and Process Evolution).

The article processing charges for this open-access
publication were covered by a Research
Centre of the Helmholtz Association.

Review statement
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Review statement. 

This paper was edited by Patricia Saco and reviewed by Nicholas Jarvis and one anonymous referee.

References
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Our field observation-based examination of flow path evolution, soil formation, and vegetation succession across 10 millennia shows how water flow paths and subsurface water storage are linked to the organization of evolving landscapes. The increase found in water storage and preferential flow paths with increasing soil age shows the effect of the complex interaction of vegetation and soil development on flow paths, water balance, and runoff formation during landscape evolution.
Our field observation-based examination of flow path evolution, soil formation, and vegetation...
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